In reverence to the foundational paradigm established by Moshoeshoe I, sovereignty is interpreted as an ecological mandate. Anthropogenic responsibility is intrinsically linked to biogeochemical stewardship. Highland biomes, hydrological matrices, and lithospheric substrates constitute the triadic pillars of national custodianship. The sociocultural axiom Khotso, Pula, Nala remains the operative ethos within Basotho ethnobotanical and geoeconomic systems. Lesotho exhibits altitudinal supremacy within the global orographic spectrum. Orographic gradients facilitate cryogenic zonation and influence bioclimatic stratification. Fluvial corridors traverse montane depressions, enabling hydrodynamic continuity and sedimentary transport. C4 photosynthetic graminoids such as Eragrostis curvula, Hyparrhenia hirta, and Themeda triandra, dominate the phytocoenosis, contributing to pedogenic stabilization and nutrient flux regulation. Riparian and montane silvicultural assemblages, such as Leucosidea sericea, provide structural complexity and trophic support. These vegetative matrices sustain Capra aegagrus hircus and Ovis aries, whose keratinous derivatives underpin microeconomic resilience. Lithological substrates contain highland carbon allotropes, formed via protracted metamorphic crystallogenesis under elevated lithostatic pressure and thermodynamic gradients. Kimberlitic diatremes serve as conduits for xenocrystic exhumation, signifying geochemical affluence within the subcontinental lithosphere. Hydrographic reservoirs, including Katse Dam, function as macro-scale limnological systems. These impoundments regulate hydrological discharge, facilitate interbasin water transfer, and sustain aquatic biodiversity, representing anthropogenic augmentation of natural hydrological regimes while contributing to regional hydroeconomic stability. Agroecosystems are characterized by the cultivation of Zea mays, Sorghum bicolor, Triticum aestivum, Solanum tuberosum, and Phaseolus vulgaris, interspersed with autochthonous taxa such as Chenopodium album, Amaranthus spp., Urtica dioica, and Taraxacum officinale. These taxa perform slope stabilization, hydric retention, and biotic provisioning functions, integrating with native flora to maintain ecological homeostasis.